1 muscle fiber contains:
Several nuclei
Several myofibrils
Myofibrils are made up of:
Sarcomeres.
Sarcomeres are made up of:
Actin filaments and myosin
filaments that slide together
and apart to create basic
muscle contraction.
Connective tissue
There are 3 layers
of connective tissue:
Epimysium--surrounds
entire muscle
Perimysium--surrounds
muscle fascicles
Endomysium--surrounds
individual muscle fibers
But how do muscle fibers
actually contract?
Cross bridges are formed between the thin actin myofilaments and thick myosin myofilaments, causing the myofilaments to slide between one another. This happens when an electrical signal comes from the motor neuron associated with the muscle and causes depolarization in the muscle fiber, which releases calcium. The excess calcium binds with troponin and this signals the actin and myosin to bond.
number of cross-bridges formed depends on abundance of actin and myosin and frequency of stimulus
Different Types of
Muscle Architecure:
Parallel and Pennate
Parallel Muscle Architecure
Parallel muscles are always
composed of relatively long fibers that
attach to tendons at the ends of the
muscles.
Subtypes of Parallel:
Strap: parallel arch. with
less prominent tendons,
so less prominent taper
at ends. Ex.--sartorius
Fusiform: parallel arch. with
more prominent tendons at
each end, so very prominent
tapers.
Pennate Muscle Architecture
Pennate muscles are always composed
of relatively short fibers that insert
obliquely into a tendon running
throughout the muscle.
Subtypes of Pennate:
Unipennate: pennate arch.
in which one tendon
penetrates the muscle.
Bipennate: pennate arch.
in which two tendons
penetrate the muscle.
Multipennate: pennate arch.
in which three or more
tendons penetrate the muscle.
Compare and Contrast: Traits of
Parallel and Pennate Muscles
Amount of contraction possible
Parallel muscles can shorten much
more than pennate muscles because
longer muscle fibers create greater
contraction compared to total muscle
size.
Angle at which muscle
fibers run
Parallel fibers run parallel
to the longitudinal muscle
axis, while pennate fibers
run oblique to the longitudinal
muscle axis because they
insert into the tendon
within the pennate muscle.
Length of muscle fibers
Parallel muscles have much
longer fibers than pennate
muscles do, because parallel
fibers run the entire length
of the muscle to attach to
tendons at each end.
Size of joint excursion
Parallel muscles can usually
create a larger joint excursion
than a pennate muscle can
because of their longer fibers.
However, this also depends on
the muscle's moment arm.
PCSA
PCSA=physiological cross-section area.
Pennate muscles have more fibers and
a larger PCSA than parallel muscles, and
this makes them capable of generating
larger forces than parallel muscles of
same size.
Muscle strength is
measured by the equation:
M=r x F
M=the moment generated
r=the muscle's moment arm
F=the muscle's contractile force
5 factors that influence a
muscle's contractile force (F)
Muscle size.
The more fibers, the larger maximum force of contraction.
This is because of the muscle's PCSA (physiological cross-sectional area)
PCSA responds to changes in activity level overtime
Stretch of the muscle.
DEFINITION: whether the muscle is shorter or longer than its optimal length.
The muscle's length is influenced by its elastic connective components.
The muscles length influences the interaction
of sarcomeres.
A muscle that is stretched too long or too
short cannot contract effectively.
Level of muscle fiber recruitment.
DEFINITION: how many muscle motor units are active.
Electromyogram (EMG) reflects the number of active motor units and frequency of their activity. As muscle's mechanical advantage increases, EMG decreases
In isometric (neither shortening, nor lengthening) contractions, number of motor units active is directly proportional to the force generated.
In con/eccentric contractions, the effect of number of motor units active depends on the mechanical advantage of that specific muscle.
Contraction velocity
DEFINITION: how fast the muscle contracts.
In concentric (shortening) contractions--speed of contraction is inversely proportional to force.
In eccentric (lengthening) contractions--speed of
contraction is directly proportional to force. Produce more F than isometric and concentric. Increases with the speed of contraction
Motor unit types composing the muscle.
Type I motor units--slow twitch.
Generate less force, but do not fatigue.
Type IIa motor units--moderate twitch.
Generate faster, tire more easily.
Type IIb motor units--fast twitch.
Generate fastest, tire most easily.
Relationship of moment arm
to muscle moment generated
The length of the moment arm is directly
proportional to the moment generated.
Greatest moment arm--when muscle's
angle of application is 90 degrees.
Least moment arm--when muscle is at
its most lengthened position.
5 things that alter muscle
function over time
Prolonged lengthening
Effect: protein synthesis and production
of additional sarcomeres, creating
hypertrophy (muscle growth).
Prolonged shortening
Effect: loss of sarcomeres,
creating atrophy (muscle loss). Transition toward type II muscle fibers
Resistance exercise
Effect: hypertrophy, increase in cross-sectional
area of muscles, protein synthesis showing
type II fibers transforming to type I.
Decreased activity
Effect: atrophy, reduction in CSA
of muscles, type I fibers begin to
transfor to type II.
Aging
Effect: loss in total strength,
fewer type II fibers, increased atrophy.
Organization of Muscle
From Oatis Chapter 4
Illustrations of Muscle Types
Relationship of moment
arm to joint excursion
The length of the muscle's moment arm
and the angular joint excursion are
inversely proportional.
Example: contraction of a muscle with a shorter
moment arm produces a larger angular excursion
than the same amount of contraction in a muscle
with a longer moment arm.
IMPORTANT DEFINITIONS TO NOTE FIRST
Moment arm of muscle: the perpendicular distance
between the muscle and the point of rotation.
Angle of application: angle between the line of pull of
a muscle and the limb to which the muscle attaches.
Anatomical moment arm of muscle: the distance along the bone between the muscle's attachment and the center of rotation of the joint. Related to true moment arm by the sine of the angle of application.
from human anatomy diagrams
Factors that Influence Muscle's Ability to Produce Motion
length of fibers composing the muscle
function of the architecture
length of muscle's moment arm
Comparison of eccentric, isometric and concentric muscle strengths with elbow positions from Oatis, Chapter 4